Chapter 4. Absorption by Roots

Means of Transport

* Diffusion, facilitated diffusion and active transport are three means of transport
in plants.
* Diffusion is a passive process; it requires no expenditure of energy.
* Diffusion facilitated by proteins without any energy expenditure is referred to as facilitated transport.
* Unlike diffusion, active transport requires energy to move molecules against the concentration quotient.
* Based on direction, short- distance transport in plants can be differentiated into three types- symport, antiport and uniport.

Imbibition and Water Potential


* Seeds absorb water by special type of diffusion called imbibitions, where solids
absorb water, causing an increase in their volume.
* The water potential gradient comprises two main components: solute potential and pressure potential.
* Pure water has the greatest possible water potential, whereas the water potential of all solutes is lower than that of pure water.
* The magnitude of the lowering of water potential due to the addition of solute is called solute potential.
* Pressure potential, which is denoted by ψp, is usually positive.

Osmosis


* The cell wall is freely permeable whereas the cell membrane is a semipermeable membrane.

* The diffusion of water across a semi- permeable membrane like the cell membrane is referred to as osmosis.
 * This potential of water molecules to move from a hypotonic solution to a hypertonic solution across a semi- permeable membrane is called osomotic potential.
* The external pressure required to prevent the water from diffusing is known as osmotic pressure.
* Numerically, osmotic pressure is equal to osmotic potential. However, their signs differ.

Plasmolysis


* It is the solution that surrounds a plant cell that decides the direction of the
water movement.
* The surrounding solution can be categorised into three types: isotonic, hypotonic and hypertonic.
* When a cell is placed in a hypertonic solution and water moves cell out of the plant cell, the cell membrane shrinks away from the cell wall. This phenomenon is called plasmolysis.
* When cells are placed in a hypertonic solution, cytoplasm builds turgor pressure against the cell wall.
* If the cell is placed in an isotonic solution, there is no net flow of water from either inside or outside the cell. The cells are said to be flaccid.

How Do Plants Absorb Water


* Root hairs present on root tips absorb both water and minerals.

* After water is absorbed, it moves deeper into the root layers by either the apoplast pathway or symplast pathway.
* The apoplast pathway is a continuous system of adjacent cell walls in a plant, interrupted only by waxy, suberised casparian strips in the root endodermis.
* The symplastic pathway is a system of interconnected protoplasms.
* Some plants have additional structures, like mycorrhiza, which help their roots in absorption.

Upward Water Movement in a Plant


* The upward movement of water in plants is made possible by a combination of
root pressure and transpiration pull.
* When mineral ions from the soil are also pushed into the root vascular tissues along with water, the pressure is known as root pressure.
* Transpiration pull is generated by loss of water from leaf stomata via transpiration.
* As Cohension- tension helps in the transpiration pull, it is also known as cohesion- tension- transpiration pull model of water transport.

Transport of Mineral

* Minerals can be absorbed from the soil both passively and by active transport.
* Plants can also remobilise minerals from older, dying parts to young, developing parts where they are required.
* Xylem and Phloem exchange small amounts of the minerals they carry.
* Minerals are transported inside plants via Xylem and Phloem elements.

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